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The e locus is the position of a gene on a chromosome with three known alleles: recessive red, ember and wild type. It was once thought that orojo may have been an allele at this locus, but it has since been discovered to be caused by the unrelated grizzly gene.

The e locus
Recessive Red Carneau pigeon
Recessive Red Carneau pigeon e//e likely plus other darkening modifiers.
Mutation type Colour modifier
Alleles recessive red, ember, wild type.